Mimulus

 Phrymaceae (Scrophulariaceae)

©The World Botanical Associates Web Page
Prepared by Richard W. Spjut
May 2004;  August 2006

Taxonomic Note: Various shrub types along the Pacific coast from Curry Co. Oregon to Isla Cedros off the coast of Baja California in Mexico, and also variants that extend inland along the inner coast ranges of California, and along the Sierra Nevada of California to the Sierra Juarez of Baja California, are treated in the Jepson Manual (Hickman 1993) under a single species, Mimulus aurantiacus.  Other authorities have in the past recognized this obvious polymorphic species to comprise 7–15 species, which have been segregated into another genus, Diplacus.  Flowers vary in color— from red to yellow—and in size, while the plants also vary in habit and in pubescence.  Although putative hybrids or intermediates between the various taxonomic entities have been viewed as justification for treatment under a single species,  it is also difficult to ignore the gross differences that are encountered in the field, especially when they appear correlated with different ecological environments  Therefore, more than one species in this  complex is recognized below.  Differences are based on the keys provided in Abrams (Illus. Fl.  Pac. States, Vol. 3, 1951, Munz & Keck (Calif. Fl.,1959), and Wiggins (Fl Baja Calif., 1981).  The type for M. aurantiacus was reportedly collected from near San Francisco or Monterrey.   A summary of the taxonomy with references is provided by Montalvo and McMillan (Oct 2003) under the USDA Forest Service International Programs website;  http://www.fs.fed.us/global/iitf/pdf/.  A  poster summary by Tulig and Clark shows the geographical occurrences of the various taxa and their intermediates or hybrids, http://www.csupomona.edu/~jcclark/pdf/p2000aspt.pdf#search=%22Mimulus%20aurantiacus%20taxonomy%22

 

Mimulus (Diplacus) aurantiacus
Humboldt Co. CA: coastal bluffs above beach near Trinidad, July 2006

 

Mimulus longiflorus
ssp. caylcinus (Diplacus calycinus)
Kern River Canyon, CA
Apr 2005

Mimulus longiflorus
Transverse Ranges, CA
Spjut 14791, Apr 2002

 

Mimulus (Diplacus) puniceus
Peninsular Ranges,
near Valley Center, CA
Photo by Susan Spjut, June 2006

Mimulus (Diplacus) puniceus
Peninsular Ranges, CA
Spjut & Marin 14663, Apr 2002

 

Mimulus (Diplacus) puniceus
Peninsular Ranges, CA
Spjut & Marin 14663, Apr 2002

 

 

Mimulus bigelovii
Mesquite Mts., CA
Spjut 15890, Mar 2005

 

Mimulus bigelovii
Mesquite Mts., CA
Spjut 15294, May 2003

 

 

Mimulus guttatus
Peninsular Ranges, CA
Near Valley Center
Photo by Susan Spjut
June 2006

 

Mimulus primuloides
 
var. pilosellus
Marble Mts. Wilderness, CA
Log Lake, July 1970

 

Mimulus primuloides
 
var. pilosellus
Marble Mts. Wilderness, CA
Clear Lake, July 1990

 

Phillips W. R, N. J. Baj,  A. A. Gunatilaka and D. G. Kingston.  1996.  C-geranyl compounds from Mimulus clevelandii. J. Nat. Prod. 59: 495–497. "Fractionation of the MeCOEt extract of Mimulus clevelandii yielded the novel 4-geranyl-5-hydroxy-2(3H)-benzofuranone (1) and the five known 6-geranylflavanones diplacone (2a), 3‘-O-methyldiplacone (2b), diplacol (2c), mimulone (2d), and 3‘-O-methyldiplacol (2e). 2D-NMR methods required revision of assignments for diplacone and diplacol and resolved the uncertainty in the site of methylation for the methyl ethers."